Viral-mediated knockdown of Atxn2 attenuates TDP-43 pathology and muscle dysfunction in the PFN1C71G ALS mouse model

Abstract Amyotrophic lateral sclerosis (ALS) is a fatal neurodegenerative disorder characterized by progressive motor neuron loss and muscle atrophy. Hyperphosphorylated aggregation of the RNA-binding protein, TDP-43, in the motor cortex and spinal cord are defining molecular features of ALS, sugges...

Full description

Saved in:
Bibliographic Details
Main Authors: Zachary C. E. Hawley, Xueying Li, Dora Bodnar, Yuanzheng Gu, Yi Luo, Daniel Ferretti, Adam Sheehy, Rachelle Driscoll, Maria I. Zavodszky, Shaolong Cao, Isabel Isaza, Luke Jandreski, Yuqing Liu, Thomas Carlile, Shih-Ching Lo, Anna Grimard, Shawn Bourque, Aditya Utturkar, Samantha Desmarais, H. Moore Arnold, Dann Huh, Edward Guilmette, Deborah Y. Kwon
Format: Article
Language:English
Published: BMC 2025-05-01
Series:Acta Neuropathologica Communications
Online Access:https://doi.org/10.1186/s40478-025-02005-z
Tags: Add Tag
No Tags, Be the first to tag this record!
_version_ 1850268601672531968
author Zachary C. E. Hawley
Xueying Li
Dora Bodnar
Yuanzheng Gu
Yi Luo
Daniel Ferretti
Adam Sheehy
Rachelle Driscoll
Maria I. Zavodszky
Shaolong Cao
Isabel Isaza
Luke Jandreski
Yuqing Liu
Thomas Carlile
Shih-Ching Lo
Anna Grimard
Shawn Bourque
Aditya Utturkar
Samantha Desmarais
H. Moore Arnold
Dann Huh
Edward Guilmette
Deborah Y. Kwon
author_facet Zachary C. E. Hawley
Xueying Li
Dora Bodnar
Yuanzheng Gu
Yi Luo
Daniel Ferretti
Adam Sheehy
Rachelle Driscoll
Maria I. Zavodszky
Shaolong Cao
Isabel Isaza
Luke Jandreski
Yuqing Liu
Thomas Carlile
Shih-Ching Lo
Anna Grimard
Shawn Bourque
Aditya Utturkar
Samantha Desmarais
H. Moore Arnold
Dann Huh
Edward Guilmette
Deborah Y. Kwon
author_sort Zachary C. E. Hawley
collection DOAJ
description Abstract Amyotrophic lateral sclerosis (ALS) is a fatal neurodegenerative disorder characterized by progressive motor neuron loss and muscle atrophy. Hyperphosphorylated aggregation of the RNA-binding protein, TDP-43, in the motor cortex and spinal cord are defining molecular features of ALS, suggesting TDP-43 dysfunction underlies disease pathogenesis. This phenomenon, however, has been difficult to recapitulate endogenously in animal models, impeding characterization of TDP-43 pathobiology in neurodegeneration. In this study, we report age-dependent accumulation of TDP-43 pathology in the spinal cord and progressive muscle-related deficits in transgenic mice expressing the ALS-associated PFN1C71G mutant protein. We show that transgenic neuronal expression of PFN1C71G induces early hyperphosphorylation of endogenous TDP-43 in the spinal cord that augments over time, preceding accumulation of insoluble non-phosphorylated TDP-43 and the manifestation of muscle denervation and motor dysfunction. Sustained knockdown of Atxn2 in the central nervous system (CNS) in pre-symptomatic PFN1C71G mice by AAV-driven expression of an artificial microRNA (AAV-amiR-Atxn2) reduces aberrant TDP-43 in the spinal cord, while delaying neurodegeneration and improving muscle and motor function. RNA-sequencing analysis of spinal cord samples from PFN1C71G mice and ALS donors show shared patterns of transcriptional perturbation, including a pro-inflammatory gene signature that is attenuated by AAV-amiR-Atxn2. Notably, impaired regulation of the PFN1C71G skeletal muscle transcriptome exceeds that of the spinal cord and is also improved by Atxn2 reduction in the CNS. Lastly, we find significant gene co-expression network homology between PFN1C71G mice and human ALS, with shared dysregulation of modules related to neuroinflammation and neuronal function and uncover novel hub genes that provide biological insight into ALS and potential drug targets that can be further investigated in this mouse model.
format Article
id doaj-art-fa19f4fbede144cd8d59b1e1cd3df8a0
institution OA Journals
issn 2051-5960
language English
publishDate 2025-05-01
publisher BMC
record_format Article
series Acta Neuropathologica Communications
spelling doaj-art-fa19f4fbede144cd8d59b1e1cd3df8a02025-08-20T01:53:25ZengBMCActa Neuropathologica Communications2051-59602025-05-0113112210.1186/s40478-025-02005-zViral-mediated knockdown of Atxn2 attenuates TDP-43 pathology and muscle dysfunction in the PFN1C71G ALS mouse modelZachary C. E. Hawley0Xueying Li1Dora Bodnar2Yuanzheng Gu3Yi Luo4Daniel Ferretti5Adam Sheehy6Rachelle Driscoll7Maria I. Zavodszky8Shaolong Cao9Isabel Isaza10Luke Jandreski11Yuqing Liu12Thomas Carlile13Shih-Ching Lo14Anna Grimard15Shawn Bourque16Aditya Utturkar17Samantha Desmarais18H. Moore Arnold19Dann Huh20Edward Guilmette21Deborah Y. Kwon22BiogenBiogenBiogenBiogenBiogenBiogenBiogenBiogenBiogenBiogenBiogenBiogenBiogenBiogenBiogenBiogenBiogenBiogenBiogenBiogenBiogenBiogenBiogenAbstract Amyotrophic lateral sclerosis (ALS) is a fatal neurodegenerative disorder characterized by progressive motor neuron loss and muscle atrophy. Hyperphosphorylated aggregation of the RNA-binding protein, TDP-43, in the motor cortex and spinal cord are defining molecular features of ALS, suggesting TDP-43 dysfunction underlies disease pathogenesis. This phenomenon, however, has been difficult to recapitulate endogenously in animal models, impeding characterization of TDP-43 pathobiology in neurodegeneration. In this study, we report age-dependent accumulation of TDP-43 pathology in the spinal cord and progressive muscle-related deficits in transgenic mice expressing the ALS-associated PFN1C71G mutant protein. We show that transgenic neuronal expression of PFN1C71G induces early hyperphosphorylation of endogenous TDP-43 in the spinal cord that augments over time, preceding accumulation of insoluble non-phosphorylated TDP-43 and the manifestation of muscle denervation and motor dysfunction. Sustained knockdown of Atxn2 in the central nervous system (CNS) in pre-symptomatic PFN1C71G mice by AAV-driven expression of an artificial microRNA (AAV-amiR-Atxn2) reduces aberrant TDP-43 in the spinal cord, while delaying neurodegeneration and improving muscle and motor function. RNA-sequencing analysis of spinal cord samples from PFN1C71G mice and ALS donors show shared patterns of transcriptional perturbation, including a pro-inflammatory gene signature that is attenuated by AAV-amiR-Atxn2. Notably, impaired regulation of the PFN1C71G skeletal muscle transcriptome exceeds that of the spinal cord and is also improved by Atxn2 reduction in the CNS. Lastly, we find significant gene co-expression network homology between PFN1C71G mice and human ALS, with shared dysregulation of modules related to neuroinflammation and neuronal function and uncover novel hub genes that provide biological insight into ALS and potential drug targets that can be further investigated in this mouse model.https://doi.org/10.1186/s40478-025-02005-z
spellingShingle Zachary C. E. Hawley
Xueying Li
Dora Bodnar
Yuanzheng Gu
Yi Luo
Daniel Ferretti
Adam Sheehy
Rachelle Driscoll
Maria I. Zavodszky
Shaolong Cao
Isabel Isaza
Luke Jandreski
Yuqing Liu
Thomas Carlile
Shih-Ching Lo
Anna Grimard
Shawn Bourque
Aditya Utturkar
Samantha Desmarais
H. Moore Arnold
Dann Huh
Edward Guilmette
Deborah Y. Kwon
Viral-mediated knockdown of Atxn2 attenuates TDP-43 pathology and muscle dysfunction in the PFN1C71G ALS mouse model
Acta Neuropathologica Communications
title Viral-mediated knockdown of Atxn2 attenuates TDP-43 pathology and muscle dysfunction in the PFN1C71G ALS mouse model
title_full Viral-mediated knockdown of Atxn2 attenuates TDP-43 pathology and muscle dysfunction in the PFN1C71G ALS mouse model
title_fullStr Viral-mediated knockdown of Atxn2 attenuates TDP-43 pathology and muscle dysfunction in the PFN1C71G ALS mouse model
title_full_unstemmed Viral-mediated knockdown of Atxn2 attenuates TDP-43 pathology and muscle dysfunction in the PFN1C71G ALS mouse model
title_short Viral-mediated knockdown of Atxn2 attenuates TDP-43 pathology and muscle dysfunction in the PFN1C71G ALS mouse model
title_sort viral mediated knockdown of atxn2 attenuates tdp 43 pathology and muscle dysfunction in the pfn1c71g als mouse model
url https://doi.org/10.1186/s40478-025-02005-z
work_keys_str_mv AT zacharycehawley viralmediatedknockdownofatxn2attenuatestdp43pathologyandmuscledysfunctioninthepfn1c71galsmousemodel
AT xueyingli viralmediatedknockdownofatxn2attenuatestdp43pathologyandmuscledysfunctioninthepfn1c71galsmousemodel
AT dorabodnar viralmediatedknockdownofatxn2attenuatestdp43pathologyandmuscledysfunctioninthepfn1c71galsmousemodel
AT yuanzhenggu viralmediatedknockdownofatxn2attenuatestdp43pathologyandmuscledysfunctioninthepfn1c71galsmousemodel
AT yiluo viralmediatedknockdownofatxn2attenuatestdp43pathologyandmuscledysfunctioninthepfn1c71galsmousemodel
AT danielferretti viralmediatedknockdownofatxn2attenuatestdp43pathologyandmuscledysfunctioninthepfn1c71galsmousemodel
AT adamsheehy viralmediatedknockdownofatxn2attenuatestdp43pathologyandmuscledysfunctioninthepfn1c71galsmousemodel
AT rachelledriscoll viralmediatedknockdownofatxn2attenuatestdp43pathologyandmuscledysfunctioninthepfn1c71galsmousemodel
AT mariaizavodszky viralmediatedknockdownofatxn2attenuatestdp43pathologyandmuscledysfunctioninthepfn1c71galsmousemodel
AT shaolongcao viralmediatedknockdownofatxn2attenuatestdp43pathologyandmuscledysfunctioninthepfn1c71galsmousemodel
AT isabelisaza viralmediatedknockdownofatxn2attenuatestdp43pathologyandmuscledysfunctioninthepfn1c71galsmousemodel
AT lukejandreski viralmediatedknockdownofatxn2attenuatestdp43pathologyandmuscledysfunctioninthepfn1c71galsmousemodel
AT yuqingliu viralmediatedknockdownofatxn2attenuatestdp43pathologyandmuscledysfunctioninthepfn1c71galsmousemodel
AT thomascarlile viralmediatedknockdownofatxn2attenuatestdp43pathologyandmuscledysfunctioninthepfn1c71galsmousemodel
AT shihchinglo viralmediatedknockdownofatxn2attenuatestdp43pathologyandmuscledysfunctioninthepfn1c71galsmousemodel
AT annagrimard viralmediatedknockdownofatxn2attenuatestdp43pathologyandmuscledysfunctioninthepfn1c71galsmousemodel
AT shawnbourque viralmediatedknockdownofatxn2attenuatestdp43pathologyandmuscledysfunctioninthepfn1c71galsmousemodel
AT adityautturkar viralmediatedknockdownofatxn2attenuatestdp43pathologyandmuscledysfunctioninthepfn1c71galsmousemodel
AT samanthadesmarais viralmediatedknockdownofatxn2attenuatestdp43pathologyandmuscledysfunctioninthepfn1c71galsmousemodel
AT hmoorearnold viralmediatedknockdownofatxn2attenuatestdp43pathologyandmuscledysfunctioninthepfn1c71galsmousemodel
AT dannhuh viralmediatedknockdownofatxn2attenuatestdp43pathologyandmuscledysfunctioninthepfn1c71galsmousemodel
AT edwardguilmette viralmediatedknockdownofatxn2attenuatestdp43pathologyandmuscledysfunctioninthepfn1c71galsmousemodel
AT deborahykwon viralmediatedknockdownofatxn2attenuatestdp43pathologyandmuscledysfunctioninthepfn1c71galsmousemodel