The LARP6 La module from Tetrabaena socialis reveals structural and functional differences from plant and animal LARP6 homologues

This study identified the LARP6 La Module from Tetrabaena socialis (T. socialis), a four-celled green algae, in an effort to better understand the evolution of LARP6 structure and RNA-binding activity in multicellular eukaryotes. Using a combination of sequence alignments, domain boundary screens, a...

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Main Authors: Emily M. Lewis, Olga Becker, Alexis N. Symons, Cora LaCoss, A. Jasmine Baclig, Avery Guzman, Charles Sanders, Leticia Gonzalez, Lisa R. Warner, Karen A. Lewis
Format: Article
Language:English
Published: Taylor & Francis Group 2025-12-01
Series:RNA Biology
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Online Access:https://www.tandfonline.com/doi/10.1080/15476286.2025.2489303
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author Emily M. Lewis
Olga Becker
Alexis N. Symons
Cora LaCoss
A. Jasmine Baclig
Avery Guzman
Charles Sanders
Leticia Gonzalez
Lisa R. Warner
Karen A. Lewis
author_facet Emily M. Lewis
Olga Becker
Alexis N. Symons
Cora LaCoss
A. Jasmine Baclig
Avery Guzman
Charles Sanders
Leticia Gonzalez
Lisa R. Warner
Karen A. Lewis
author_sort Emily M. Lewis
collection DOAJ
description This study identified the LARP6 La Module from Tetrabaena socialis (T. socialis), a four-celled green algae, in an effort to better understand the evolution of LARP6 structure and RNA-binding activity in multicellular eukaryotes. Using a combination of sequence alignments, domain boundary screens, and structural modelling, we recombinantly expressed and isolated the TsLARP6 La Module to > 98% purity for in vitro biochemical characterization. The La Module is stably folded and exerts minimal RNA binding activity against single-stranded homopolymeric RNAs. Surprisingly, it exhibits low micromolar binding affinity for the vertebrate LARP6 cognate ligand, a bulged-stem loop found in the 5’UTR of collagen type I mRNA, but does not bind double-stranded RNAs of similar size. These result suggests that the TsLARP6 La Module may prefer structured RNA ligands. In contrast, however, the TsLARP6 La Module does not exhibit the RNA chaperone activity that is observed in vertebrate homologs. Therefore, we conclude that protist LARP6 may have both distinct RNA ligands and binding mechanisms from the previously characterized LARP6 proteins of animals and vascular plants, thus establishing a distinct third class of the LARP6 protein family.
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spelling doaj-art-e9e49fb3f05a4b718d20c87c69f856272025-08-20T02:08:35ZengTaylor & Francis GroupRNA Biology1547-62861555-85842025-12-012211910.1080/15476286.2025.2489303The LARP6 La module from Tetrabaena socialis reveals structural and functional differences from plant and animal LARP6 homologuesEmily M. Lewis0Olga Becker1Alexis N. Symons2Cora LaCoss3A. Jasmine Baclig4Avery Guzman5Charles Sanders6Leticia Gonzalez7Lisa R. Warner8Karen A. Lewis9Department of Chemistry and Biochemistry, Texas State University, San Marcos, TX, USADepartment of Chemistry and Biochemistry, Texas State University, San Marcos, TX, USADepartment of Chemistry and Biochemistry, Texas State University, San Marcos, TX, USADepartment of Chemistry and Biochemistry, Texas State University, San Marcos, TX, USADepartment of Chemistry and Biochemistry, Boise State University, Boise, ID, USADepartment of Chemistry and Biochemistry, Texas State University, San Marcos, TX, USADepartment of Chemistry and Biochemistry, Texas State University, San Marcos, TX, USADepartment of Chemistry and Biochemistry, Texas State University, San Marcos, TX, USADepartment of Chemistry and Biochemistry, Boise State University, Boise, ID, USADepartment of Chemistry and Biochemistry, Texas State University, San Marcos, TX, USAThis study identified the LARP6 La Module from Tetrabaena socialis (T. socialis), a four-celled green algae, in an effort to better understand the evolution of LARP6 structure and RNA-binding activity in multicellular eukaryotes. Using a combination of sequence alignments, domain boundary screens, and structural modelling, we recombinantly expressed and isolated the TsLARP6 La Module to > 98% purity for in vitro biochemical characterization. The La Module is stably folded and exerts minimal RNA binding activity against single-stranded homopolymeric RNAs. Surprisingly, it exhibits low micromolar binding affinity for the vertebrate LARP6 cognate ligand, a bulged-stem loop found in the 5’UTR of collagen type I mRNA, but does not bind double-stranded RNAs of similar size. These result suggests that the TsLARP6 La Module may prefer structured RNA ligands. In contrast, however, the TsLARP6 La Module does not exhibit the RNA chaperone activity that is observed in vertebrate homologs. Therefore, we conclude that protist LARP6 may have both distinct RNA ligands and binding mechanisms from the previously characterized LARP6 proteins of animals and vascular plants, thus establishing a distinct third class of the LARP6 protein family.https://www.tandfonline.com/doi/10.1080/15476286.2025.2489303La-related proteinLARPLARP6RNA binding proteinalgaeprotist
spellingShingle Emily M. Lewis
Olga Becker
Alexis N. Symons
Cora LaCoss
A. Jasmine Baclig
Avery Guzman
Charles Sanders
Leticia Gonzalez
Lisa R. Warner
Karen A. Lewis
The LARP6 La module from Tetrabaena socialis reveals structural and functional differences from plant and animal LARP6 homologues
RNA Biology
La-related protein
LARP
LARP6
RNA binding protein
algae
protist
title The LARP6 La module from Tetrabaena socialis reveals structural and functional differences from plant and animal LARP6 homologues
title_full The LARP6 La module from Tetrabaena socialis reveals structural and functional differences from plant and animal LARP6 homologues
title_fullStr The LARP6 La module from Tetrabaena socialis reveals structural and functional differences from plant and animal LARP6 homologues
title_full_unstemmed The LARP6 La module from Tetrabaena socialis reveals structural and functional differences from plant and animal LARP6 homologues
title_short The LARP6 La module from Tetrabaena socialis reveals structural and functional differences from plant and animal LARP6 homologues
title_sort larp6 la module from tetrabaena socialis reveals structural and functional differences from plant and animal larp6 homologues
topic La-related protein
LARP
LARP6
RNA binding protein
algae
protist
url https://www.tandfonline.com/doi/10.1080/15476286.2025.2489303
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