A morphological cell atlas of the freshwater sponge Ephydatia muelleri with key insights from targeted single-cell transcriptomes

Abstract How animal cell types, tissues, and regional body plans arose is a fundamental question in EvoDevo. Many current efforts attempt to link genetic information to the morphology of cells, tissues and regionalization of animal body plans using single-cell sequencing of cell populations. However...

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Main Authors: Sally P. Leys, Lauren Grombacher, Daniel Field, Glen R. D. Elliott, Vanessa R. Ho, Amanda S. Kahn, Pamela J. Reid, Ana Riesgo, Emilio Lanna, Yuriy Bobkov, Joseph F. Ryan, April L. Horton
Format: Article
Language:English
Published: BMC 2025-02-01
Series:EvoDevo
Online Access:https://doi.org/10.1186/s13227-025-00237-7
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author Sally P. Leys
Lauren Grombacher
Daniel Field
Glen R. D. Elliott
Vanessa R. Ho
Amanda S. Kahn
Pamela J. Reid
Ana Riesgo
Emilio Lanna
Yuriy Bobkov
Joseph F. Ryan
April L. Horton
author_facet Sally P. Leys
Lauren Grombacher
Daniel Field
Glen R. D. Elliott
Vanessa R. Ho
Amanda S. Kahn
Pamela J. Reid
Ana Riesgo
Emilio Lanna
Yuriy Bobkov
Joseph F. Ryan
April L. Horton
author_sort Sally P. Leys
collection DOAJ
description Abstract How animal cell types, tissues, and regional body plans arose is a fundamental question in EvoDevo. Many current efforts attempt to link genetic information to the morphology of cells, tissues and regionalization of animal body plans using single-cell sequencing of cell populations. However, a lack of in-depth understanding of the morphology of non-bilaterian animals remains a considerable block to understanding the transitions between bilaterian and non-bilaterian cells and tissues. Sponges (Porifera), one of the earliest diverging animal phyla, pose a particular challenge to this endeavour, because their body plans lack mouths, gut, conventional muscle and nervous systems. With a goal to help bridge this gap, we have studied the morphology, behaviour and transcriptomics of cells and tissue types of an easily accessible and well-studied species of freshwater sponge, Ephydatia muelleri. New features described here include: a polarized external epithelium, a new contractile sieve cell that forms the entry to incurrent canals, motile cilia on apopyle cells at the exit of choanocyte chambers, and non-motile cilia on cells in excurrent canals and oscula. Imaging cells in vivo shows distinct behavioural characteristics of motile cells in the mesohyl. Transcriptomic phenotypes of three cell types (cystencytes, choanocytes and archaeocytes) captured live indicate that cell-type transcriptomes are distinct. Importantly, individual archaeocytes show a range of transcriptomic phenotypes which is supported by the distinct expression of different genes by subsets of this cell type. In contrast, all five choanocyte cells sampled live revealed highly uniform transcriptomes with significantly fewer genes expressed than in other cell types. Our study shows that sponges have tissues whose morphology and cell diversity are both functionally complex, but which together enable the sponge, like other metazoans, to sense and respond to stimuli.
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spelling doaj-art-e899e0f4741d4231b714fc93dcfbca4b2025-08-20T03:00:59ZengBMCEvoDevo2041-91392025-02-0116112610.1186/s13227-025-00237-7A morphological cell atlas of the freshwater sponge Ephydatia muelleri with key insights from targeted single-cell transcriptomesSally P. Leys0Lauren Grombacher1Daniel Field2Glen R. D. Elliott3Vanessa R. Ho4Amanda S. Kahn5Pamela J. Reid6Ana Riesgo7Emilio Lanna8Yuriy Bobkov9Joseph F. Ryan10April L. Horton11Department of Biological Sciences, University of AlbertaDepartment of Biological Sciences, University of AlbertaDepartment of Biological Sciences, University of AlbertaDepartment of Biological Sciences, University of AlbertaDepartment of Biological Sciences, University of AlbertaDepartment of Biological Sciences, University of AlbertaDepartment of Biological Sciences, University of AlbertaDepartment of Biological Sciences, University of AlbertaDepartment of Biological Sciences, University of AlbertaWhitney Laboratory for Marine Bioscience, University of FloridaWhitney Laboratory for Marine Bioscience, University of FloridaBiology Department, Bates CollegeAbstract How animal cell types, tissues, and regional body plans arose is a fundamental question in EvoDevo. Many current efforts attempt to link genetic information to the morphology of cells, tissues and regionalization of animal body plans using single-cell sequencing of cell populations. However, a lack of in-depth understanding of the morphology of non-bilaterian animals remains a considerable block to understanding the transitions between bilaterian and non-bilaterian cells and tissues. Sponges (Porifera), one of the earliest diverging animal phyla, pose a particular challenge to this endeavour, because their body plans lack mouths, gut, conventional muscle and nervous systems. With a goal to help bridge this gap, we have studied the morphology, behaviour and transcriptomics of cells and tissue types of an easily accessible and well-studied species of freshwater sponge, Ephydatia muelleri. New features described here include: a polarized external epithelium, a new contractile sieve cell that forms the entry to incurrent canals, motile cilia on apopyle cells at the exit of choanocyte chambers, and non-motile cilia on cells in excurrent canals and oscula. Imaging cells in vivo shows distinct behavioural characteristics of motile cells in the mesohyl. Transcriptomic phenotypes of three cell types (cystencytes, choanocytes and archaeocytes) captured live indicate that cell-type transcriptomes are distinct. Importantly, individual archaeocytes show a range of transcriptomic phenotypes which is supported by the distinct expression of different genes by subsets of this cell type. In contrast, all five choanocyte cells sampled live revealed highly uniform transcriptomes with significantly fewer genes expressed than in other cell types. Our study shows that sponges have tissues whose morphology and cell diversity are both functionally complex, but which together enable the sponge, like other metazoans, to sense and respond to stimuli.https://doi.org/10.1186/s13227-025-00237-7
spellingShingle Sally P. Leys
Lauren Grombacher
Daniel Field
Glen R. D. Elliott
Vanessa R. Ho
Amanda S. Kahn
Pamela J. Reid
Ana Riesgo
Emilio Lanna
Yuriy Bobkov
Joseph F. Ryan
April L. Horton
A morphological cell atlas of the freshwater sponge Ephydatia muelleri with key insights from targeted single-cell transcriptomes
EvoDevo
title A morphological cell atlas of the freshwater sponge Ephydatia muelleri with key insights from targeted single-cell transcriptomes
title_full A morphological cell atlas of the freshwater sponge Ephydatia muelleri with key insights from targeted single-cell transcriptomes
title_fullStr A morphological cell atlas of the freshwater sponge Ephydatia muelleri with key insights from targeted single-cell transcriptomes
title_full_unstemmed A morphological cell atlas of the freshwater sponge Ephydatia muelleri with key insights from targeted single-cell transcriptomes
title_short A morphological cell atlas of the freshwater sponge Ephydatia muelleri with key insights from targeted single-cell transcriptomes
title_sort morphological cell atlas of the freshwater sponge ephydatia muelleri with key insights from targeted single cell transcriptomes
url https://doi.org/10.1186/s13227-025-00237-7
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