A naturally occurring SNP modulates thermotolerance divergence among grapevines

Abstract With the increasing challenges posed by global warming and climate change, heat stress has become a significant threat to the sustainable production of grapevines. However, the genetic basis of grapevine thermotolerance remains poorly understood. Here, we combine genome-wide association stu...

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Main Authors: Haiyang Chen, Haibo Yu, Ling Yuan, Lingchao Kong, Shenchang Li, Xiongjun Cao, Yang Li, Yi Wang, Ling Lin, Rongrong Guo, Taili Xie, Wei Duan, Zhanwu Dai, Peige Fan, Shaohua Li, Zhenchang Liang, Lijun Wang
Format: Article
Language:English
Published: Nature Portfolio 2025-06-01
Series:Nature Communications
Online Access:https://doi.org/10.1038/s41467-025-60209-2
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Summary:Abstract With the increasing challenges posed by global warming and climate change, heat stress has become a significant threat to the sustainable production of grapevines. However, the genetic basis of grapevine thermotolerance remains poorly understood. Here, we combine genome-wide association study with transcriptomic profiling to identify TTC4 (thermotolerance on chromosome 4), a gene encoding a WRKY transcription factor, as a key determinant of thermotolerance in grapevine. TTC4 directly activates two thermotolerance-related genes, HSP18.1 and APX3. We also identify a heat-suppressed repressor SPL13 (SQUAMOSA-promoter binding protein-like 13) that cannot bind to the GTAT element (TTC4 T(7631)) in intron 2 of TTC4, but can bind to the natural variant, GTAC (TTC4 C(7631)). Grapevine accessions with TTC4 C/C(7631) genotype exhibit significantly lower thermotolerance compared to those with the TTC4 T/T(7631) and TTC4 C/T(7631) genotypes. This fine-tuned regulation contributes to thermotolerance divergence among grapevine populations. The TTC4 T(7631) haplotype holds significant potential as a genetic resource for breeding thermotolerant grapevine varieties.
ISSN:2041-1723