Genomic DNA barcodes provide novel insights into species delimitation in the complex Camellia sect. Thea (Theaceae)

Abstract Background Species delimitation within Camellia sect. Thea is taxonomically challenging due to its complex evolutionary history. This study aims to utilize nuclear and chloroplast data as genomic DNA barcodes to delimit species within this economically important group of plants. Results Who...

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Main Authors: Zongfang Shen, Yujing Feng, Michael Möller, Kevin S. Burgess, Hantao Qin, Junbo Yang, Zhiqiong Mo, Hongtao Li, Dezhu Li, Lianming Gao
Format: Article
Language:English
Published: BMC 2025-05-01
Series:BMC Plant Biology
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Online Access:https://doi.org/10.1186/s12870-025-06612-9
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author Zongfang Shen
Yujing Feng
Michael Möller
Kevin S. Burgess
Hantao Qin
Junbo Yang
Zhiqiong Mo
Hongtao Li
Dezhu Li
Lianming Gao
author_facet Zongfang Shen
Yujing Feng
Michael Möller
Kevin S. Burgess
Hantao Qin
Junbo Yang
Zhiqiong Mo
Hongtao Li
Dezhu Li
Lianming Gao
author_sort Zongfang Shen
collection DOAJ
description Abstract Background Species delimitation within Camellia sect. Thea is taxonomically challenging due to its complex evolutionary history. This study aims to utilize nuclear and chloroplast data as genomic DNA barcodes to delimit species within this economically important group of plants. Results Whole genome sequencing (WGS) data were obtained for 98 accessions representing all but one species in C. sect. Thea. Based on 759 high-quality SCNs, 98 whole chloroplast genomes, and by using 2× coverage clean reads from WGS for Skmer analyses, we found that combining the findings from these three data sets resulted in nearly complete species delimitation and resolution of all interspecific relationships within C. sect. Thea. We also found support for the taxonomic elevation of two varieties (C. sinensis var. assamica and C. tachangensis var. remotiserrata) to species status (C. assamica and C. remotiserrata, respectively). Furthermore, we confirmed that C. formosensis represents a distinct species. Gene tree discordances, chloroplast-nuclear conflicts and complex network-like phylogenetic relationships were observed in C. sect. Thea. Conclusion Compared with the use of single parentally inherited chloroplast data sources, utilizing both uniparentally inherited chloroplast data and biparentally inherited nuclear data improved the species delimitation of taxa within C. sect. Thea. The intricate phylogenetic relationships observed are likely a result of widespread past hybridization and chloroplast capture events among species within this group, which may have blurred the species boundaries. Our novel approach to species delimitation within C. sect. Thea may serve as a blueprint for employing genomic DNA barcodes in other taxa with complex histories, and will significantly contribute to the conservation of cultivated tea plant species and their wild relatives.
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spelling doaj-art-a570b294a7fc45c8b65f320a6ce71bdd2025-08-20T02:55:38ZengBMCBMC Plant Biology1471-22292025-05-0125111310.1186/s12870-025-06612-9Genomic DNA barcodes provide novel insights into species delimitation in the complex Camellia sect. Thea (Theaceae)Zongfang Shen0Yujing Feng1Michael Möller2Kevin S. Burgess3Hantao Qin4Junbo Yang5Zhiqiong Mo6Hongtao Li7Dezhu Li8Lianming Gao9Germplasm Bank of Wild Species, Kunming Institute of Botany, Chinese Academy of SciencesGermplasm Bank of Wild Species, Kunming Institute of Botany, Chinese Academy of SciencesRoyal Botanic Garden EdinburghDepartment of Biomedical Sciences, Mercer University School of MedicineGermplasm Bank of Wild Species, Kunming Institute of Botany, Chinese Academy of SciencesGermplasm Bank of Wild Species, Kunming Institute of Botany, Chinese Academy of SciencesGermplasm Bank of Wild Species, Kunming Institute of Botany, Chinese Academy of SciencesGermplasm Bank of Wild Species, Kunming Institute of Botany, Chinese Academy of SciencesGermplasm Bank of Wild Species, Kunming Institute of Botany, Chinese Academy of SciencesGermplasm Bank of Wild Species, Kunming Institute of Botany, Chinese Academy of SciencesAbstract Background Species delimitation within Camellia sect. Thea is taxonomically challenging due to its complex evolutionary history. This study aims to utilize nuclear and chloroplast data as genomic DNA barcodes to delimit species within this economically important group of plants. Results Whole genome sequencing (WGS) data were obtained for 98 accessions representing all but one species in C. sect. Thea. Based on 759 high-quality SCNs, 98 whole chloroplast genomes, and by using 2× coverage clean reads from WGS for Skmer analyses, we found that combining the findings from these three data sets resulted in nearly complete species delimitation and resolution of all interspecific relationships within C. sect. Thea. We also found support for the taxonomic elevation of two varieties (C. sinensis var. assamica and C. tachangensis var. remotiserrata) to species status (C. assamica and C. remotiserrata, respectively). Furthermore, we confirmed that C. formosensis represents a distinct species. Gene tree discordances, chloroplast-nuclear conflicts and complex network-like phylogenetic relationships were observed in C. sect. Thea. Conclusion Compared with the use of single parentally inherited chloroplast data sources, utilizing both uniparentally inherited chloroplast data and biparentally inherited nuclear data improved the species delimitation of taxa within C. sect. Thea. The intricate phylogenetic relationships observed are likely a result of widespread past hybridization and chloroplast capture events among species within this group, which may have blurred the species boundaries. Our novel approach to species delimitation within C. sect. Thea may serve as a blueprint for employing genomic DNA barcodes in other taxa with complex histories, and will significantly contribute to the conservation of cultivated tea plant species and their wild relatives.https://doi.org/10.1186/s12870-025-06612-9Camellia sect. TheaSpecies delimitationGenomic DNA barcodesChloroplast genomesSingle-copy nuclear orthologous genes
spellingShingle Zongfang Shen
Yujing Feng
Michael Möller
Kevin S. Burgess
Hantao Qin
Junbo Yang
Zhiqiong Mo
Hongtao Li
Dezhu Li
Lianming Gao
Genomic DNA barcodes provide novel insights into species delimitation in the complex Camellia sect. Thea (Theaceae)
BMC Plant Biology
Camellia sect. Thea
Species delimitation
Genomic DNA barcodes
Chloroplast genomes
Single-copy nuclear orthologous genes
title Genomic DNA barcodes provide novel insights into species delimitation in the complex Camellia sect. Thea (Theaceae)
title_full Genomic DNA barcodes provide novel insights into species delimitation in the complex Camellia sect. Thea (Theaceae)
title_fullStr Genomic DNA barcodes provide novel insights into species delimitation in the complex Camellia sect. Thea (Theaceae)
title_full_unstemmed Genomic DNA barcodes provide novel insights into species delimitation in the complex Camellia sect. Thea (Theaceae)
title_short Genomic DNA barcodes provide novel insights into species delimitation in the complex Camellia sect. Thea (Theaceae)
title_sort genomic dna barcodes provide novel insights into species delimitation in the complex camellia sect thea theaceae
topic Camellia sect. Thea
Species delimitation
Genomic DNA barcodes
Chloroplast genomes
Single-copy nuclear orthologous genes
url https://doi.org/10.1186/s12870-025-06612-9
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