Food reward entrainment increases mealtime anxiety in goldfish via a ghrelin-dependent mechanism

Abstract Food anticipatory activity (FAA), the increase in locomotor activity prior to a scheduled feeding time, is linked not only to energy demands but also to food reward expectations. However, the mechanisms behind FAA remain unclear. We hypothesize that FAA induces anxiety due to reward anticip...

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Main Authors: Lisbeth Herrera-Castillo, Nuria Saiz, Nuria de Pedro, Esther Isorna
Format: Article
Language:English
Published: Nature Portfolio 2025-07-01
Series:Scientific Reports
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Online Access:https://doi.org/10.1038/s41598-025-13194-x
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author Lisbeth Herrera-Castillo
Nuria Saiz
Nuria de Pedro
Esther Isorna
author_facet Lisbeth Herrera-Castillo
Nuria Saiz
Nuria de Pedro
Esther Isorna
author_sort Lisbeth Herrera-Castillo
collection DOAJ
description Abstract Food anticipatory activity (FAA), the increase in locomotor activity prior to a scheduled feeding time, is linked not only to energy demands but also to food reward expectations. However, the mechanisms behind FAA remain unclear. We hypothesize that FAA induces anxiety due to reward anticipation. To test this, anxiety-like behavior was assessed in goldfish using open field and black-and-white preference tests under different feeding conditions and daytimes. Increased thigmotactic and scototactic behavior during FAA, compared to the post-FAA period, was observed, even in animals under constant light. Additionally, an unexpected meal did not reduce anxiety, while more hours of fasting did not increase it, suggesting that FAA is an anxiogenic state driven by a food-entrained oscillator rather than energy status. Ghrelin signaling was further explored using two antagonists, both of which reduced anxiety-like behavior when administered during FAA. Moreover, ghrelin administration induced anxiety-like behavior that was reversed by co-injection with an antagonist, supporting its anxiogenic role. These findings suggest a shared neural mechanism between FAA and anxiety, mediated by ghrelin. The possible involvement of ghrelin in activating dopamine-reward circuits in teleosts should be explored in future studies to gain further insight into the neurobiological basis of food anticipatory activity.
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spelling doaj-art-9aca9f49a49a40e4aaec00c3f9d9bdba2025-08-20T03:42:41ZengNature PortfolioScientific Reports2045-23222025-07-0115111510.1038/s41598-025-13194-xFood reward entrainment increases mealtime anxiety in goldfish via a ghrelin-dependent mechanismLisbeth Herrera-Castillo0Nuria Saiz1Nuria de Pedro2Esther Isorna3Department of Genetics, Physiology and Microbiology, Faculty of Biological Sciences, Complutense University of MadridDepartment of Genetics, Physiology and Microbiology, Faculty of Biological Sciences, Complutense University of MadridDepartment of Genetics, Physiology and Microbiology, Faculty of Biological Sciences, Complutense University of MadridDepartment of Genetics, Physiology and Microbiology, Faculty of Biological Sciences, Complutense University of MadridAbstract Food anticipatory activity (FAA), the increase in locomotor activity prior to a scheduled feeding time, is linked not only to energy demands but also to food reward expectations. However, the mechanisms behind FAA remain unclear. We hypothesize that FAA induces anxiety due to reward anticipation. To test this, anxiety-like behavior was assessed in goldfish using open field and black-and-white preference tests under different feeding conditions and daytimes. Increased thigmotactic and scototactic behavior during FAA, compared to the post-FAA period, was observed, even in animals under constant light. Additionally, an unexpected meal did not reduce anxiety, while more hours of fasting did not increase it, suggesting that FAA is an anxiogenic state driven by a food-entrained oscillator rather than energy status. Ghrelin signaling was further explored using two antagonists, both of which reduced anxiety-like behavior when administered during FAA. Moreover, ghrelin administration induced anxiety-like behavior that was reversed by co-injection with an antagonist, supporting its anxiogenic role. These findings suggest a shared neural mechanism between FAA and anxiety, mediated by ghrelin. The possible involvement of ghrelin in activating dopamine-reward circuits in teleosts should be explored in future studies to gain further insight into the neurobiological basis of food anticipatory activity.https://doi.org/10.1038/s41598-025-13194-xFeeding entrained oscillator (FEO)TeleostsFood anticipatory activity (FAA)Food seekingGhrelinHedonic system
spellingShingle Lisbeth Herrera-Castillo
Nuria Saiz
Nuria de Pedro
Esther Isorna
Food reward entrainment increases mealtime anxiety in goldfish via a ghrelin-dependent mechanism
Scientific Reports
Feeding entrained oscillator (FEO)
Teleosts
Food anticipatory activity (FAA)
Food seeking
Ghrelin
Hedonic system
title Food reward entrainment increases mealtime anxiety in goldfish via a ghrelin-dependent mechanism
title_full Food reward entrainment increases mealtime anxiety in goldfish via a ghrelin-dependent mechanism
title_fullStr Food reward entrainment increases mealtime anxiety in goldfish via a ghrelin-dependent mechanism
title_full_unstemmed Food reward entrainment increases mealtime anxiety in goldfish via a ghrelin-dependent mechanism
title_short Food reward entrainment increases mealtime anxiety in goldfish via a ghrelin-dependent mechanism
title_sort food reward entrainment increases mealtime anxiety in goldfish via a ghrelin dependent mechanism
topic Feeding entrained oscillator (FEO)
Teleosts
Food anticipatory activity (FAA)
Food seeking
Ghrelin
Hedonic system
url https://doi.org/10.1038/s41598-025-13194-x
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