The role of ABC transporter DrrABC in the export of PDIM in Mycobacterium tuberculosis

The Mycobacterium tuberculosis virulence lipid phthiocerol dimycocerosate (PDIM) is exported by a complex mechanism that involves multiple proteins including the Resistance-Nodulation-Division (RND) transporter MmpL7 and the lipoprotein LppX. Here, we probe the role of the putative heterooligomeric...

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Main Authors: Nabiela Moolla, Helen Weaver, Rebeca Bailo, Albel Singh, Vassiliy N. Bavro, Apoorva Bhatt
Format: Article
Language:English
Published: Elsevier 2024-12-01
Series:The Cell Surface
Online Access:http://www.sciencedirect.com/science/article/pii/S2468233024000148
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author Nabiela Moolla
Helen Weaver
Rebeca Bailo
Albel Singh
Vassiliy N. Bavro
Apoorva Bhatt
author_facet Nabiela Moolla
Helen Weaver
Rebeca Bailo
Albel Singh
Vassiliy N. Bavro
Apoorva Bhatt
author_sort Nabiela Moolla
collection DOAJ
description The Mycobacterium tuberculosis virulence lipid phthiocerol dimycocerosate (PDIM) is exported by a complex mechanism that involves multiple proteins including the Resistance-Nodulation-Division (RND) transporter MmpL7 and the lipoprotein LppX. Here, we probe the role of the putative heterooligomeric ATP-Binding Cassette (ABC) transporter complex composed of DrrA, DrrB and DrrC in PDIM transport by constructing a set of individual null mutants of drrA, drrB and drrC in the vaccine strain Mycobacterium bovis BCG. Loss of all three, or individual drr genes, all resulted in a complete loss of PDIM export to the outer envelope of the mycobacterial cell. Furthermore, guided by a bioinformatic analysis we interrogated specific signature residues within the DrrABC to demonstrate that it is indeed an ABC transporter, and our modelling, together with the mutagenesis identify it as a member of the Type V family of ABC exporters. We identify several unique structural elements of the transporter, including a non-canonical C-terminally inserted domain (CTD) structure within DrrA, which may account for its functional properties.
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issn 2468-2330
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publishDate 2024-12-01
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series The Cell Surface
spelling doaj-art-6dee0c938b9a4a4cbb2d52de9f4451c82025-08-20T02:50:20ZengElsevierThe Cell Surface2468-23302024-12-011210013210.1016/j.tcsw.2024.100132The role of ABC transporter DrrABC in the export of PDIM in Mycobacterium tuberculosisNabiela Moolla0Helen Weaver1Rebeca Bailo2Albel Singh3Vassiliy N. Bavro4Apoorva Bhatt5School of Biosciences and Institute of Microbiology and Infection, University of Birmingham, Edgbaston, Birmingham B15 2TT, United KingdomSchool of Biosciences and Institute of Microbiology and Infection, University of Birmingham, Edgbaston, Birmingham B15 2TT, United KingdomSchool of Biosciences and Institute of Microbiology and Infection, University of Birmingham, Edgbaston, Birmingham B15 2TT, United KingdomSchool of Biosciences and Institute of Microbiology and Infection, University of Birmingham, Edgbaston, Birmingham B15 2TT, United KingdomSchool of Life Sciences, University of Essex, Colchester CO4 3SQ, United Kingdom; Corresponding authors.School of Biosciences and Institute of Microbiology and Infection, University of Birmingham, Edgbaston, Birmingham B15 2TT, United Kingdom; Corresponding authors.The Mycobacterium tuberculosis virulence lipid phthiocerol dimycocerosate (PDIM) is exported by a complex mechanism that involves multiple proteins including the Resistance-Nodulation-Division (RND) transporter MmpL7 and the lipoprotein LppX. Here, we probe the role of the putative heterooligomeric ATP-Binding Cassette (ABC) transporter complex composed of DrrA, DrrB and DrrC in PDIM transport by constructing a set of individual null mutants of drrA, drrB and drrC in the vaccine strain Mycobacterium bovis BCG. Loss of all three, or individual drr genes, all resulted in a complete loss of PDIM export to the outer envelope of the mycobacterial cell. Furthermore, guided by a bioinformatic analysis we interrogated specific signature residues within the DrrABC to demonstrate that it is indeed an ABC transporter, and our modelling, together with the mutagenesis identify it as a member of the Type V family of ABC exporters. We identify several unique structural elements of the transporter, including a non-canonical C-terminally inserted domain (CTD) structure within DrrA, which may account for its functional properties.http://www.sciencedirect.com/science/article/pii/S2468233024000148
spellingShingle Nabiela Moolla
Helen Weaver
Rebeca Bailo
Albel Singh
Vassiliy N. Bavro
Apoorva Bhatt
The role of ABC transporter DrrABC in the export of PDIM in Mycobacterium tuberculosis
The Cell Surface
title The role of ABC transporter DrrABC in the export of PDIM in Mycobacterium tuberculosis
title_full The role of ABC transporter DrrABC in the export of PDIM in Mycobacterium tuberculosis
title_fullStr The role of ABC transporter DrrABC in the export of PDIM in Mycobacterium tuberculosis
title_full_unstemmed The role of ABC transporter DrrABC in the export of PDIM in Mycobacterium tuberculosis
title_short The role of ABC transporter DrrABC in the export of PDIM in Mycobacterium tuberculosis
title_sort role of abc transporter drrabc in the export of pdim in mycobacterium tuberculosis
url http://www.sciencedirect.com/science/article/pii/S2468233024000148
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