Geographical distribution of Cryptococcus neoformans in Iran, pathogenic factors, and antifungal susceptibility profile

Abstract Background Approximately 70 species of Cryptococcus have been identified, some of which are human and animal pathogens, while others are environmental saprophytes. Cryptococcus neoformans is one of the most important pathogenic species, and pigeon droppings serve as the most important ecolo...

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Main Authors: Sara Hamzehee, Marzieh Halvaeizadeh, Maral Gharaghani, Hadis Jafarian, Simin Taghipour, Zahra Seifi, Sahar Hivary, Sadegh Noripour-Sisakht, Ali Zarei-Mahmoudabadi
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Language:English
Published: BMC 2025-08-01
Series:BMC Microbiology
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Online Access:https://doi.org/10.1186/s12866-025-04292-x
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author Sara Hamzehee
Marzieh Halvaeizadeh
Maral Gharaghani
Hadis Jafarian
Simin Taghipour
Zahra Seifi
Sahar Hivary
Sadegh Noripour-Sisakht
Ali Zarei-Mahmoudabadi
author_facet Sara Hamzehee
Marzieh Halvaeizadeh
Maral Gharaghani
Hadis Jafarian
Simin Taghipour
Zahra Seifi
Sahar Hivary
Sadegh Noripour-Sisakht
Ali Zarei-Mahmoudabadi
author_sort Sara Hamzehee
collection DOAJ
description Abstract Background Approximately 70 species of Cryptococcus have been identified, some of which are human and animal pathogens, while others are environmental saprophytes. Cryptococcus neoformans is one of the most important pathogenic species, and pigeon droppings serve as the most important ecological niche. This study aimed to investigate the frequency of C. neoformans in pigeon droppings from various Iranian provinces, as well as assess the pathogenic factors and antifungal susceptibility of the isolates. Methods 428 pigeon droppings samples were collected from 10 Iranian provinces and cultured on Niger seed agar. DNAs were extracted, and CNLAC1 (Outer and Inner) and STR1 primers were used for the detection of species and varieties of C. neoformans, respectively. Virulence factors were also identified using specific culture media under appropriate conditions. Antifungal susceptibility testing was conducted against 8 antifungals using CLSI guidelines. Results Out of the 428 examined samples, 37 (8.6%) were positive for C. neoformans. All isolates exhibited capsule production and dark brown pigmentation as key pathogenic factors. Urease and catalase activities were present in all isolates. Based on the established ECVs guidelines, all isolates displayed wild-type phenotypes against voriconazole and 5-flucytosine. While 100% isolates were non-wild type against amphotericin B. Conclusions The frequency of C. neoformans was 8.6% in Iranian cities, with a high (31.5%) frequency in Kermanshah and a low (3.1%) frequency in Fars province. All isolates showed sensitivity to voriconazole and 5-flucytosine. Additionally, pathogenic factors were present in all strains to varying degrees.
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spelling doaj-art-5cb940da04ae47d7afbf43f4fa43ba862025-08-24T11:10:48ZengBMCBMC Microbiology1471-21802025-08-012511910.1186/s12866-025-04292-xGeographical distribution of Cryptococcus neoformans in Iran, pathogenic factors, and antifungal susceptibility profileSara Hamzehee0Marzieh Halvaeizadeh1Maral Gharaghani2Hadis Jafarian3Simin Taghipour4Zahra Seifi5Sahar Hivary6Sadegh Noripour-Sisakht7Ali Zarei-Mahmoudabadi8Department of Medical Mycology, School of Medicine, Ahvaz Jundishapur University of Medical SciencesDepartment of Medical Mycology, School of Medicine, Ahvaz Jundishapur University of Medical SciencesMedicinal Plants Research Center, Yasuj University of Medical SciencesProfessor Alborzi Clinical Microbiology Research Center, Shiraz University of Medical SciencesDepartment of Medical Parasitology and Mycology, Faculty of Medicine, Shahrekord University of Medical SciencesLaboratory Sciences Research Center, Department of Medical Parasitology and Mycology, Faculty of Paramedicine, Golestan University of Medical SciencesShahid Jahangirzadeh ClinicMedicinal Plants Research Center, Yasuj University of Medical SciencesDepartment of Medical Mycology, School of Medicine, Ahvaz Jundishapur University of Medical SciencesAbstract Background Approximately 70 species of Cryptococcus have been identified, some of which are human and animal pathogens, while others are environmental saprophytes. Cryptococcus neoformans is one of the most important pathogenic species, and pigeon droppings serve as the most important ecological niche. This study aimed to investigate the frequency of C. neoformans in pigeon droppings from various Iranian provinces, as well as assess the pathogenic factors and antifungal susceptibility of the isolates. Methods 428 pigeon droppings samples were collected from 10 Iranian provinces and cultured on Niger seed agar. DNAs were extracted, and CNLAC1 (Outer and Inner) and STR1 primers were used for the detection of species and varieties of C. neoformans, respectively. Virulence factors were also identified using specific culture media under appropriate conditions. Antifungal susceptibility testing was conducted against 8 antifungals using CLSI guidelines. Results Out of the 428 examined samples, 37 (8.6%) were positive for C. neoformans. All isolates exhibited capsule production and dark brown pigmentation as key pathogenic factors. Urease and catalase activities were present in all isolates. Based on the established ECVs guidelines, all isolates displayed wild-type phenotypes against voriconazole and 5-flucytosine. While 100% isolates were non-wild type against amphotericin B. Conclusions The frequency of C. neoformans was 8.6% in Iranian cities, with a high (31.5%) frequency in Kermanshah and a low (3.1%) frequency in Fars province. All isolates showed sensitivity to voriconazole and 5-flucytosine. Additionally, pathogenic factors were present in all strains to varying degrees.https://doi.org/10.1186/s12866-025-04292-xCryptococcus neoformansVirulence factorsAntifungal susceptibilityCapsuleMelaninIran
spellingShingle Sara Hamzehee
Marzieh Halvaeizadeh
Maral Gharaghani
Hadis Jafarian
Simin Taghipour
Zahra Seifi
Sahar Hivary
Sadegh Noripour-Sisakht
Ali Zarei-Mahmoudabadi
Geographical distribution of Cryptococcus neoformans in Iran, pathogenic factors, and antifungal susceptibility profile
BMC Microbiology
Cryptococcus neoformans
Virulence factors
Antifungal susceptibility
Capsule
Melanin
Iran
title Geographical distribution of Cryptococcus neoformans in Iran, pathogenic factors, and antifungal susceptibility profile
title_full Geographical distribution of Cryptococcus neoformans in Iran, pathogenic factors, and antifungal susceptibility profile
title_fullStr Geographical distribution of Cryptococcus neoformans in Iran, pathogenic factors, and antifungal susceptibility profile
title_full_unstemmed Geographical distribution of Cryptococcus neoformans in Iran, pathogenic factors, and antifungal susceptibility profile
title_short Geographical distribution of Cryptococcus neoformans in Iran, pathogenic factors, and antifungal susceptibility profile
title_sort geographical distribution of cryptococcus neoformans in iran pathogenic factors and antifungal susceptibility profile
topic Cryptococcus neoformans
Virulence factors
Antifungal susceptibility
Capsule
Melanin
Iran
url https://doi.org/10.1186/s12866-025-04292-x
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